Bordered Screw-moss vs clouded brindle

Tortula marginata compared with Apamea epomidion

Key Differences

Bordered Screw-moss is Endangered while clouded brindle is Least Concern.

Taxonomic Classification

Rank	Bordered Screw-moss	clouded brindle
Kingdom	Plantae (Plants)	Animalia (Animals)
Phylum	Bryophyta	Arthropoda (Arthropods)
Class	Bryopsida (Bryopsida)	Insecta (Insects)
Order	Pottiales (Pottiales)	Lepidoptera (Butterflies & Moths)
Family	Pottiaceae	Noctuidae
Genus	Tortula	Apamea
Species	Tortula marginata	Apamea epomidion

Conservation Status

Bordered Screw-moss

EN — Endangered

clouded brindle

LC — Least Concern

Physical Characteristics

Attribute	Bordered Screw-moss	clouded brindle
Diet	—	—
Average Lifespan	—	—
Average Length	—	—
Average Weight	—	—

Habitat & Geographic Range

Bordered Screw-moss

Habitat

Found across multiple habitat types including temperate broadleaf and mixed forests, tundra, and tropical and subtropical moist broadleaf forests spanning the Australasia and Oceanian realms.

Range

Distributed across Belgium, Luxembourg, New Zealand, Norway, and Portugal. Currently classified as Endangered on the IUCN Red List, this species faces significant conservation challenges across its range.

clouded brindle

Habitat

Typically found in virtually all terrestrial and freshwater habitats.

Range

Distributed across Belgium, Denmark, Norway, and Sweden.

Bordered Screw-moss

The Bordered Screw-moss (Tortula marginata) is a species in the genus Tortula. It is currently classified as Endangered on the IUCN Red List. Found across multiple habitat types including temperate broadleaf and mixed forests, tundra, and tropical and subtropical moist broadleaf forests spanning the Australasia and Oceanian realms.

clouded brindle

The clouded brindle (Apamea epomidion) is a noctuid moth in the family Noctuidae found across temperate Europe and extending into western Asia. The adult wingspan measures approximately 35–45 mm with typical brindle-patterned forewings in grey-brown and buff tones with subtle cross-lines and stigmata characteristic of the Apamea genus. The term 'clouded' refers to diffuse cloud-like darker shading areas across the forewing surface. Adults fly in one generation from June to August, attracted to light and flowers at night. The larvae are internal feeders within grass stems and roots, feeding on coarse grass species such as Brachypodium sylvaticum and Deschampsia in woodland rides, scrub margins, and rough grassland habitats. The pupal stage overwinters in soil or within plant debris. The clouded brindle inhabits structurally diverse woodland edge habitats with a mixture of tall grasses, scrub, and open canopy woodland rides that provide both larval foodplants and adult resting sites. Changes in woodland management, particularly reduction of coppicing and shading of woodland rides, may affect this and related grass-feeding brindle moth species.